Nature

A fascinating profile of a 33-year-old silverback named Titus, the leader of a gorilla clan in the mountainous region between Rwanda and Congo who faces a challenge for supremacy from his second-in-command, Kuryama. Archival footage and the observations of researchers fill in his backstory, including how, as a young adult, he engaged in secret liaisons with females behind the back of pack leader Beetsme, then led a rare bloodless coup against Beetsme.